Hi Dave,
Great picture of what looks to be a tidy cockpit. Yes: separate pitch-levers - maybe the Heron was to be unique in combining pitch control with throttle.
It appears that, like the post-war Heron (and Dove?), the applied pressure of the wheel brakes was commanded by variable displacement of the captain's thumb-control on his/her spectacles. The differential effect for steering would be controlled with the rudder bar. I wonder if the brakes were hydraulic or (like the Heron and Dove) pneumatic. My guess would be pneumatic.
The original tail-fin config mentioned by
Stanwell is best visible on that
video at 1:13 and 1:35.