Low fuel was a tricky situation to deal with, especially if Holding (with turns always in one direction) fuel would tip from the higher to the lower wing tanks due to an always open high level weir, the type's anherdral added to the imbalance which had to be managed via cross feeding.
OK, I'll bite the bait on your hook:
Why would an aircraft in a balanced turn run fuel from the higher wing to the lower wing in such a turn? How does the fuel in the higher wing 'know' that it is higher than the fuel in the other wing?