Widger I don't think that you can say the way Harrier was operated from "carriers" would be the way that F35 would be from the QE class. The huge difference is that the Harrier GR was thrown in late on in life - with F35 it can be planned from the start. Everyone can go to F35 with an understanding of what is required from them, fundamentally different to some guy who "didn't sign up to be on a bloody boat".
At the end of the day there needs to be an assessment (I really don't want to call it a TNA) and the secret is to find the right balance of what has been done before and what new technology enables. This would then drive how much time is needed on board, in the sim or on a dummy deck etc. You then flex this depending on defence need - if we need aircraft in afghan for another 15 years then you lean carrier training as much as possible. If we are going to the Falklands which has been taken because everyone was drunk on Friday night, then you major on carrier ops (or lots and lots of tanking!).
My personal opinion is that the RN will simply have the attitude that they are Naval Power assets and should be on the carrier whatever and will be extremely reticent to let them go elsewhere. I have seen this attitude again and again with the RN - they are extremely protectionist. By contrast the RAF will simply look how best to employ Air Power in a given situation. But I am sure you will disagree.