Air Passenger Duty: Airline mark up?

[ExXB]

Fare quote

Just did a fare quote for VS LHR-JFK return and got:

Fare (A1): VS LONNYC OLXMEG fare (rules) , psgr type ADT £43.00 Fare (A2): VS NYCLON OLXMEG fare (rules) , psgr type ADT £43.00 Tax: USDA APHIS Fee $5.00 Tax: US Immigration Fee $7.00 Tax: US Customs Fee $5.50 Tax: United Kingdom Air Passengers Duty £45.00 Tax: United Kingdom Passenger Service Charge £21.20 Tax: US International Arrival Tax $16.10 Tax: VS YQ surcharge £111.00 Tax: US International Departure Tax $16.10 Tax: US September 11th Security Fee $2.50 Tax: US Passenger Facility Charge $4.50 Total for 1 adult passenger: £298.50
The YQ (fuel) surcharge (and the fare) goes to VS, the rest to governments.